Campbell Biology · Chapter 39
Plant Responses to Internal and External Signals
pp. 821–849 · 5 sections
Plants can't walk away from shade, drought or a hungry caterpillar, so they respond by changing how they grow, what genes they switch on and what chemicals they make. This chapter follows those responses from the signaling pathways inside cells to hormones, light sensing, day-length timing, gravity and touch, stress and defense against attackers. Most plant details here are outside the current AP course, but the chapter is a strong review of tested ideas: signal transduction, feedback, gene regulation and organisms responding to their environment.
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39.1 From signal to response in plant cells
pp. 821–824
Reception, transduction and response are tested in Topics 4.1–4.3, and switching genes on and off is Topic 6.5. The plant example (a seedling greening in light) is background; you won't need its specific second messengers.
In the course: Topic 4.1 Cell Communication, Topic 4.2 Introduction to Signal Transduction, Topic 4.3 Signal Transduction Pathways, Topic 6.5 Regulation of Gene Expression (notes, videos and more questions)
Key points
- A plant can't move toward food or away from trouble, so it answers changes around it mostly by adjusting how it grows and develops. Inside its cells, though, signaling follows the same three steps you learn for animal cells: reception, transduction and response.
- A seedling growing underground looks very different from one in the light. In the dark it is etiolated: a long, pale, fast-growing stem, tiny folded leaves and little root, which spends its stored food on reaching the surface. Once light hits it, de-etiolation (greening) begins: the stem slows down, the leaves open up and chlorophyll is made.
- Reception: each receptor is a protein whose shape shifts only when its own particular signal arrives. For greening, that receptor is phytochrome, a light-absorbing protein that sits in the cytoplasm instead of in the plasma membrane.
- Transduction: small second messengers, such as cyclic GMP (cGMP) and Ca²⁺, carry the message onward and amplify it, so even very faint light can trigger a big response. Protein kinases pass the signal along by adding phosphate groups, and protein phosphatases remove those phosphates so the pathway shuts off when the signal stops.
- Response: a pathway can switch on proteins the cell already has (fast, often by phosphorylation) or change which genes are transcribed (slower). Transcription factors can be activators that turn genes up or repressors that turn them down. Greening needs many new proteins, including photosynthesis enzymes, enzymes that build chlorophyll and enzymes that change hormone levels.
- Mutants are a key tool for working out a pathway. A plant missing one step may fail to respond, and a plant missing a repressor may act as though the signal is always there. Comparing such mutants shows the order of the steps.
Key terms (13)
- etiolation
- The way a seedling grows in darkness: a long, pale stem, small folded leaves and few roots, so it can push up to the light fast.
- de-etiolation (greening)
- The switch a seedling makes once it reaches light: stem growth slows, the leaves open and turn green, and roots grow.
- receptor
- A protein that recognizes one particular signal and shifts its shape when it does, which sets the cell's response in motion.
- phytochrome
- A plant photoreceptor protein that mainly senses red and far-red light. It controls greening, germination and many other responses to light.
- signal transduction pathway
- The chain of steps inside a cell that turns a received signal into a response.
- second messenger
- A small molecule or ion inside the cell, like Ca²⁺ or cGMP, that passes a signal on from the receptor and amplifies it.
- cyclic GMP (cGMP)
- A small second-messenger molecule made from GTP. In plants it helps relay light signals.
- calcium ions (Ca²⁺)
- Ions kept at very low levels in the cytosol. When channels open and the level rises, Ca²⁺ acts as a second messenger.
- protein kinase
- An enzyme that adds a phosphate group to another protein, often switching it on. Kinases pass signals along in a chain.
- protein phosphatase
- An enzyme that removes phosphate groups from proteins, helping to switch a pathway off when the signal is gone.
- transcription factor
- A protein that binds DNA and turns specific genes up (an activator) or down (a repressor).
- post-translational modification
- A change made to a protein after it is built, such as adding a phosphate group. It lets a cell switch on proteins it already has.
- amplification
- How a signal grows stronger at each step of a pathway, so a few receptor events can change thousands of molecules.
Check yourself: 39.1 From signal to response in plant cells
4 questions on 39.1 From signal to response in plant cells. Pick an answer to see if you got it, and why.
Root cells of a grass detect a molecule released by a soil fungus and, about an hour later, begin secreting a defensive enzyme. Which sequence best describes what happens inside each root cell?
Two related wildflower species have seeds holding enough stored food for about 6 days of growth in darkness. In the dark, species X's stem grows 1.5 cm per day and species Y's grows 0.4 cm per day. Assuming each stem grows straight up, from about how deep in the soil could each species' seedling still reach the light?
In a model plant signaling pathway, one activated receptor switches on 12 molecules of an enzyme, and each of those enzyme molecules then makes 400 molecules of a second messenger. How many second-messenger molecules result from one activated receptor?
Leaf cells are treated with a drug that blocks protein phosphatases. The cells then get a short pulse of a signal that works through a protein kinase pathway. Compared with untreated cells, what would most likely happen?
0 of 4 answered
39.2 Plant hormones and what they do
pp. 824–835
You won't be asked to name plant hormones or list their jobs, but ethylene in fruit ripening is a standard positive feedback example (Topic 4.4), and hormones show long-distance signaling and pathways (Topics 4.1 and 4.3).
In the course: Topic 4.1 Cell Communication, Topic 4.3 Signal Transduction Pathways, Topic 4.4 Feedback, Topic 8.1 Responses to the Environment (notes, videos and more questions)
Key points
- Plant hormones are signal molecules made in tiny amounts that change growth, development or responses to the environment, often in a different part of the plant from where they're made. One hormone can do many jobs, and many outcomes depend on the balance between two or more hormones rather than on one alone. Some biologists call them plant growth regulators.
- Hormones were first found by studying shoots that bend toward light (phototropism). Early experiments with grass seedlings showed that the tip senses light and that a chemical travels down from it to the zone that bends. That chemical was named auxin; the main natural form is indoleacetic acid (IAA). Auxin builds up on the shaded side, cells there stretch more, and the shoot bends toward the light.
- Auxin is made mostly in shoot tips and young leaves and moves downward from cell to cell in one direction (polar transport), because the carrier proteins that pump it out sit at the lower end of each cell. It speeds cell elongation by the acid growth mechanism: proton pumps acidify the cell wall, low pH activates expansins that loosen the wall, and water pressure stretches the cell. Auxin also shapes branching, leaf placement and vein patterns, and helps form roots. At high levels it slows elongation, partly by triggering ethylene.
- Cytokinins, made mainly in roots and carried up in the xylem, work with auxin to make cells divide. In tissue culture, more cytokinin than auxin gives shoots and more auxin gives roots. Cytokinins also wake side buds and slow leaf aging. Apical dominance, where a growing shoot tip keeps the buds below it dormant, depends on auxin, cytokinins and strigolactones together.
- Gibberellins (GAs) make stems elongate, sometimes into a sudden flower stalk (bolting). They also help fruit grow and end seed dormancy: in a cereal grain, the embryo releases GA and the aleurone layer answers by making enzymes such as α-amylase that digest stored starch. GA works by getting a repressor (DELLA) protein destroyed. Brassinosteroids are steroid hormones that promote elongation and cell division, and unlike animal steroids they bind a receptor on the cell surface.
- Abscisic acid (ABA) generally slows growth. It keeps seeds dormant (the ABA-to-GA balance decides whether a seed sprouts) and closes stomata during drought by making guard cells lose K⁺ and water. Strigolactones are made in roots. They hold back shoot branching, attract helpful mycorrhizal fungi, and can trigger germination of parasitic plants' seeds.
- Ethylene is a gas made during stress, ripening, aging and leaf drop. In a seedling blocked by an obstacle it causes the triple response: slower lengthening, a thicker stem and sideways growth. As a leaf ages and makes less auxin, the base of its stalk becomes sensitive to ethylene and the leaf drops. In many fruits, ethylene sets ripening going and the ripening fruit pours out still more of the gas, a positive feedback loop. Because it's a gas, it drifts to nearby fruit too, so one ripe fruit hurries the rest along.
Key terms (15)
- plant hormone
- A signal molecule made in small amounts in one part of a plant that changes growth, development or responses, often somewhere else.
- phototropism
- Growth that bends a plant organ toward light (positive) or away from it (negative), caused by cells on one side stretching more.
- auxin
- A plant hormone, mainly IAA, that makes young shoot cells stretch and helps control bending, branching, root formation and leaf patterns.
- polar transport
- Auxin's one-way, cell-to-cell flow from shoot tip toward base, caused by exit carriers sitting at the bottom end of each cell.
- acid growth hypothesis
- The idea that auxin makes cells grow by turning on proton pumps that acidify the cell wall, loosening it so water pressure can stretch the cell.
- expansin
- A wall protein switched on by low pH that loosens the links between cellulose fibers so a growing cell can stretch.
- cytokinin
- A hormone made mostly in roots and other actively growing tissues. Together with auxin it drives cell division, and it wakes side buds and delays leaf aging.
- apical dominance
- When a growing shoot tip keeps the side buds below it from growing. Removing the tip often makes a plant bushier.
- gibberellin (GA)
- A hormone that makes stems lengthen, helps fruit grow and ends seed dormancy. It works by having repressor proteins destroyed.
- brassinosteroid
- A steroid hormone in plants that boosts growth by making cells stretch and divide. Unlike animal steroids, it's detected by a receptor on the cell surface.
- abscisic acid (ABA)
- A hormone that slows growth, keeps seeds dormant and closes stomata when water is short.
- strigolactone
- A hormone made in roots that suppresses shoot branching and helps roots partner with mycorrhizal fungi.
- ethylene
- A gaseous hormone involved in fruit ripening, aging, leaf drop and responses to stress and obstacles.
- triple response
- A seedling's ethylene-driven reaction to an obstacle: slower lengthening, a thicker stem and growth sideways.
- abscission
- When a plant drops a leaf, flower or fruit by breaking down a weak layer of cells at its base. The ethylene-auxin balance controls the timing.
Check yourself: 39.2 Plant hormones and what they do
4 questions on 39.2 Plant hormones and what they do. Pick an answer to see if you got it, and why.
Young stem segments were floated for 3 hours in different solutions, and their growth was measured. Results: weak pH 7 buffer, 0.2 mm; weak pH 7 buffer + auxin, 1.4 mm; pH 4.5 buffer (no auxin), 1.3 mm; strong pH 7 buffer that stops the cell walls from turning acidic + auxin, 0.3 mm. Which conclusion do the data best support?
A thale cress seedling is lit from its left side only and begins bending to the left. Researchers then measure auxin in the left and right halves of the stem's growing zone. Which result would they most likely find?
A gall-forming soil bacterium (Agrobacterium) inserts two genes into plant cells: one for making auxin and one for making cytokinin. The infected cells keep dividing into a disorganized lump called a tumor. Researchers infect kalanchoe stems with a mutant strain whose auxin gene is broken but whose cytokinin gene still works. What would the tumors most likely do?
Gibberellin (GA) promotes stem growth by causing DELLA proteins, which repress growth genes, to be destroyed. Many high-yield 'semi-dwarf' wheat varieties carry a DELLA protein that GA can no longer mark for destruction. If these semi-dwarf plants were sprayed with extra GA, what would most likely happen?
0 of 4 answered
39.3 Sensing light, time and season
pp. 835–841
Topic 8.1 covers how organisms respond to their surroundings, and phototropism, day-length flowering and daily rhythms are classic plant cases of it. You won't need the names of the phytochrome forms or the blue-light receptors.
In the course: Topic 8.1 Responses to the Environment, Topic 4.2 Introduction to Signal Transduction, Topic 6.5 Regulation of Gene Expression (notes, videos and more questions)
Key points
- For a plant, light is information as well as energy. Plants sense whether light is present, which direction it comes from, how bright it is and what color it is. Changes in a plant's form caused by light are called photomorphogenesis.
- An action spectrum shows how well each wavelength drives a response. If it matches the absorption spectrum of a pigment, that pigment is probably the receptor. Red and blue light turn out to control most light-driven development.
- Blue-light receptors include phototropins, which control bending toward light and the movement of chloroplasts, and cryptochromes, which slow stem growth once a seedling reaches light. Blue light also helps open stomata.
- Phytochrome switches between two forms. Pr absorbs red light and turns into Pfr; Pfr absorbs far-red light and turns back into Pr. Pfr is usually the active form, and the last color a plant receives wins. Many small seeds germinate only when light turns enough phytochrome into Pfr, so they sprout where a seedling can reach light quickly.
- Leaves absorb red light but let far-red through, so under other plants the red-to-far-red ratio falls and less phytochrome is in the Pfr form. Plants that need full sun respond by growing taller and branching less, which is called shade avoidance.
- Many plant activities rise and fall on a roughly 24-hour circadian rhythm that keeps going even in constant light or dark. On its own the clock's free-running period drifts a little from 24 hours. It is built from clock genes whose proteins, after a delay, shut down their own transcription (a negative feedback loop), and light detected by phytochromes and blue-light receptors resets it each day.
- Plants track the seasons by night length (photoperiodism). What 'short-day' plants actually need is a long night: they flower when darkness lasts longer than a critical length. Long-day plants flower when it's shorter, and day-neutral plants ignore it. A brief red flash in the middle of the night breaks the night, and far-red right after cancels the flash. Leaves sense the photoperiod and send a signal (florigen, now known to be largely the FT protein) through the phloem to the shoot tip. Some plants also need a long cold spell first (vernalization).
Key terms (14)
- photomorphogenesis
- Changes in a plant's form and development that are triggered by light.
- action spectrum
- A graph of how well each wavelength of light drives a process. Matching it to a pigment's absorption reveals the receptor.
- phototropin
- A blue-light receptor that controls bending toward light and chloroplast movements.
- cryptochrome
- A blue-light receptor that helps slow stem growth when a seedling reaches light and helps set the daily clock.
- Pr and Pfr
- The two forms of the phytochrome light switch. Red light flips Pr into Pfr (usually the active form), and far-red light flips Pfr back into Pr.
- shade avoidance
- A sun-loving plant's response to a low red-to-far-red ratio: growing taller and branching less to escape neighbors' shade.
- circadian rhythm
- A cycle of about 24 hours that keeps running even without outside cues, like daily changes in gene activity or stomatal opening.
- free-running period
- The length of one cycle of a circadian rhythm when an organism is kept in constant conditions, usually a bit more or less than 24 hours.
- entrainment
- The daily resetting of the biological clock by outside cues, mainly light at dawn, so it stays matched to the 24-hour day.
- photoperiodism
- A response to how long the days and nights are, such as flowering at a set time of year.
- short-day plant
- A plant that flowers when nights are longer than a critical length, so it typically blooms after midsummer, once nights lengthen.
- long-day plant
- A plant that flowers when nights are shorter than a critical length, so it typically blooms as midsummer approaches.
- critical night length
- The number of hours of uninterrupted darkness that decides whether a plant that times flowering by day length will bloom.
- florigen
- The mobile flowering signal made in leaves. It is now known to be largely the FT protein, which travels to the shoot tip.
Check yourself: 39.3 Sensing light, time and season
4 questions on 39.3 Sensing light, time and season. Pick an answer to see if you got it, and why.
In dim light, the chloroplasts in spinach leaf cells gather along the cell walls that face the light. Researchers measured how well each color of light caused this movement (1.0 = most effective): 400 nm, 0.70; 450 nm, 1.00; 500 nm, 0.40; 550 nm, 0.05; 660 nm, 0.05; 730 nm, 0.00. Which kind of photoreceptor most likely controls the movement?
Small seeds of a meadow herb were kept moist under four conditions for 5 days. Germination: open sunlight, 88%; sunlight filtered through a layer of green leaves held above the dish, 12%; sunlight filtered through a clear plastic sheet held the same way, 85%; darkness, 9%. Which explanation fits best?
Bean plants in a growth chamber get bright white light. In one group, LED lamps add extra far-red light while the amount of red light stays the same. Compared with controls, how would the extra-far-red group most likely grow?
Plants grown on a normal day-night cycle were moved into constant dim light at hour 0. The mRNA for one gene peaked at hours 6, 31, 56 and 81. Which conclusion is best supported?
0 of 4 answered
39.4 Gravity, touch and environmental stress
pp. 841–845
The exam won't ask about statoliths or heat-shock proteins by name, but these responses to the environment (Topic 8.1) run on tested ideas: water potential (Topic 2.7), membrane fluidity (Topic 2.3) and protein denaturation (Topic 3.2).
In the course: Topic 8.1 Responses to the Environment, Topic 2.7 Tonicity and Osmoregulation, Topic 2.3 Plasma Membrane, Topic 3.2 Environmental Impacts on Enzyme Function (notes, videos and more questions)
Key points
- Gravitropism makes roots grow down (positive) and shoots grow up (negative), whichever way a seed ends up in the soil. In root cap cells, dense starch-filled plastids called statoliths sink toward the bottom of each cell, and auxin then piles up along the underside of the root. Because high auxin slows elongation in roots, the top side outgrows the bottom and the root turns downward. Plants that lack statoliths still respond, just more slowly, so gravity is probably sensed in more than one way.
- Plants are very sensitive to touch. Plants that are often shaken, brushed or blown by wind grow shorter and sturdier (thigmomorphogenesis). Tendrils coil around a support when touched because cells on the side away from the contact grow faster (thigmotropism).
- Some leaves fold within seconds of a touch. Motor cells in a swollen joint at the leaf base (a pulvinus) release K⁺, water follows by osmosis, and the cells go limp. The signal spreads as electrical action potentials, which are much slower than nerve impulses in animals.
- Drought: guard cells lose turgor and stomata close, and rising ABA keeps them closed. Leaves may roll up or drop, and roots stop growing in dry topsoil but keep growing in moister soil below. Saving water also cuts photosynthesis, which is why drought lowers crop yields.
- Flooding: waterlogged soil has little oxygen, so roots can't respire aerobically. Ethylene triggers programmed death of some cells in the root cortex, leaving air channels that carry oxygen down. Salt: salty soil has a lower water potential, so roots struggle to take up water even from wet soil, and ions like Na⁺ can poison cells. Many plants build up harmless solutes to lower their own water potential, and salt-tolerant halophytes have extra tricks such as glands that excrete salt.
- Heat: evaporation through stomata cools leaves, but closing stomata to save water gives that up. When tissues get very hot (around 40 °C for many temperate plants), cells ramp up heat-shock proteins, chaperones that help other proteins keep or recover their shape. Cold: membranes stiffen, so plants raise the share of unsaturated fatty acids to keep them fluid. Freezing: ice forms outside the cells and draws water out of them, and hardy plants store sugars and make antifreeze proteins that limit ice damage.
Key terms (13)
- gravitropism
- A growth response to gravity. Roots curve downward (positive gravitropism) and shoots curve upward (negative gravitropism).
- statolith
- A dense, starch-filled plastid that settles to the low side of certain cells, such as those in the root cap, helping the plant sense which way is down.
- thigmomorphogenesis
- Changes in a plant's form, such as shorter, thicker stems, caused by repeated touch, wind or shaking.
- thigmotropism
- Growth that bends toward or around something the plant touches, like a tendril coiling around a stick.
- pulvinus
- A swollen joint at the base of a leaf or leaflet whose motor cells change turgor to move the leaf.
- action potential (plants)
- A quick electrical signal that travels through plant tissue, triggering fast responses like leaf folding. It is far slower than a nerve impulse.
- abiotic stress
- Damage or strain caused by the nonliving surroundings, such as too little or too much water, salty soil, or extreme heat or cold.
- aerenchyma
- Root or stem tissue with large air channels, often formed by programmed cell death in low-oxygen, waterlogged soil.
- halophyte
- A plant adapted to grow in salty soil, for example by pumping salt out through special glands.
- compatible solute
- A dissolved substance, like certain sugars or amino acids, that a cell can store in large amounts without harm to lower its water potential.
- heat-shock protein
- A protein made in larger amounts at high temperature that helps other proteins keep or regain their shape.
- membrane fluidity
- How freely membrane lipids and proteins can move. More unsaturated fatty acids keep a membrane fluid in the cold.
- antifreeze protein
- A protein that binds tiny ice crystals and keeps them from growing, limiting freezing damage.
Check yourself: 39.4 Gravity, touch and environmental stress
4 questions on 39.4 Gravity, touch and environmental stress. Pick an answer to see if you got it, and why.
A seedling is laid on its side in the dark. In both its root and its shoot, auxin builds up on the lower side. Yet the root curves down and the shoot curves up. Which explanation fits best?
A student thinks roots sense gravity when dense starch-filled plastids (statoliths) in root cap cells settle toward the lower cell wall. Which observation would most strongly support this idea?
To toughen tomato transplants, a grower runs gentle fans over one bench of seedlings for three weeks so the plants sway all day, and keeps another bench in still air. Results: swayed plants, 31 cm tall with 6.1 mm stems; still-air plants, 42 cm tall with 5.2 mm stems. Which description is correct?
Each evening, the leaves of a prayer plant fold upward. The fold happens at a swollen joint at the base of the leaf (a pulvinus), where a group of 'motor' cells suddenly loses turgor. What would most likely happen if those motor cells were treated with a drug that blocks their potassium (K⁺) channels?
0 of 4 answered
39.5 Defending against herbivores and pathogens
pp. 845–847
Plant immunity isn't in the current course, but plant-herbivore and plant-pathogen battles are community interactions (Topic 8.5) shaped by natural selection (Topic 7.2), and the hypersensitive response is signaling that ends in programmed cell death (Topic 4.3).
In the course: Topic 8.5 Community Ecology, Topic 4.3 Signal Transduction Pathways, Topic 4.1 Cell Communication, Topic 7.2 Natural Selection (notes, videos and more questions)
Key points
- Plants defend against herbivores with physical barriers such as thorns, spines, tough leaves and hairs (trichomes), and with chemicals that taste bad, are toxic or block digestion. Many chemical defenses are made only after damage begins, which saves resources when no one is eating the plant.
- Chewing damage plus chemicals in the attacker's saliva switch on signaling, with jasmonic acid as a key hormone, that turns on defense genes across the plant. Damaged plants can also release airborne volatile chemicals that attract the herbivore's predators or parasites and can warn neighboring plants.
- Against pathogens, the first barrier is the plant's outer covering: the epidermis with its waxy cuticle, and bark on older stems. Viruses, bacteria and fungi get in through wounds and natural openings such as stomata.
- Once inside, pathogens face two layers of recognition. Receptors on plant cells detect molecules that many microbes share, such as pieces of bacterial flagellin, and switch on basic defenses. Successful pathogens deliver effector proteins that suppress those defenses.
- Plants fight back with resistance (R) proteins that recognize specific effectors. In gene-for-gene recognition, a plant R gene matches a pathogen gene called an avirulence (Avr) gene, so named because the pathogen fails when it's recognized. A match sets off a strong defense, and no match usually means disease.
- In the hypersensitive response, cells around the point of infection pump out antimicrobial compounds (phytoalexins) and PR proteins and toughen their walls. Then they die on purpose, leaving the pathogen stuck in a patch of dead tissue. Signals released there, such as methyl salicylate, travel through the phloem; salicylic acid then switches on defense genes plant-wide, giving broad protection for days to weeks. This is systemic acquired resistance.
- Strong defenses are expensive, so a plant usually holds them in reserve and turns them on once it senses it's under attack. Plants and their attackers keep evolving in response to each other, which is why both sides carry so many recognition and counter-recognition genes.
Key terms (14)
- herbivory
- Animals eating plants. It's a constant stress that plants defend against.
- trichome
- A hair-like outgrowth of the plant's epidermis that can block, trap or poison small herbivores.
- volatile compound
- A chemical that easily becomes a gas. Damaged plants release them to attract the herbivore's enemies or warn neighbors.
- jasmonic acid
- A plant hormone that switches on defenses, like digestion-blocking proteins, after wounding or insect attack.
- pathogen
- A virus, bacterium, fungus or other organism that causes disease.
- effector
- A protein a pathogen sends into plant cells to suppress defenses or redirect the cell's resources.
- resistance (R) gene
- A plant gene for a receptor that recognizes a specific pathogen effector and switches on strong defenses.
- avirulence (Avr) gene
- A pathogen gene for an effector that a matching R protein can recognize, which makes the pathogen fail on that plant.
- gene-for-gene recognition
- Resistance that depends on a plant R gene matching a pathogen Avr gene. No match usually means disease.
- hypersensitive response
- A local defense: cells around an infection make antimicrobial chemicals, wall themselves off and then die, so the pathogen is stuck in dead tissue.
- phytoalexin
- An antimicrobial compound a plant makes at an infection site to kill or slow bacteria and fungi.
- PR protein
- A pathogenesis-related protein made during infection. Many are enzymes that attack pathogens' cell walls.
- systemic acquired resistance
- Broad, plant-wide protection that follows a local infection, switched on by signals with salicylic acid at the center.
- salicylic acid
- A plant hormone that builds up after infection and turns on defense genes throughout the plant.
Check yourself: 39.5 Defending against herbivores and pathogens
4 questions on 39.5 Defending against herbivores and pathogens. Pick an answer to see if you got it, and why.
In gene-for-gene recognition, a plant resists a pathogen strain when one of its resistance (R) genes matches one of the strain's avirulence (Avr) genes. Plant line A has R1, line B has R2 and line C has R1 and R2. Strain X has Avr1, strain Y has Avr2 and strain Z has neither. Which pairing is most likely to result in disease?
In the hypersensitive response, leaf cells around an infection site make antimicrobial compounds, toughen their walls and then kill themselves. How does destroying its own cells help the plant?
One lower leaf of each cucumber plant is infected with a mild bacterial strain. Five days later, an upper leaf is challenged with a fungus. Average fungal lesions per upper leaf: normal plants, no first infection, 24; normal plants, first infection, 6; plants with a gene that breaks down salicylic acid, first infection, 22. Which conclusion is best supported?
In a two-choice test, 40 predatory bugs could each walk toward air from caterpillar-damaged plants or air from undamaged plants. 30 chose the damaged plants and 10 chose the undamaged plants. With a null hypothesis of no preference, the chi-square value is 10.0 (critical value 3.84 at p = 0.05, 1 degree of freedom). Which conclusion is correct?
0 of 4 answered